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Seledets. 2005. Ecological area of species: karyological aspects. Pages 95--97 in Karyology, Karyosystematics and Molecular Phylogeny. St. Petersburg, Russia.
Probatova. 2006. Further chromosome studies on vascular plant species from Sakhalin, Moneron and Kurile Islands. Biodivers. Biogeogr. Kuril Islands Sakhalin 2: 93–110.